Deseadan

The Deseadan (Spanish: Deseadense) age is a period of geologic time (29.0–21.0 Ma) within the Oligocene epoch of the Paleogene to the Early Miocene epoch of the Neogene, used more specifically within the SALMA classification of South America. It follows the Tinguirirican and precedes the Colhuehuapian age.

Etymology

The age is named after the Deseado Formation of the Deseado Massif in eastern Patagonia, Argentina.

Formations

Locations of Deseadan formations

Formation bold is type	Country	Basin	Notes
Deseado Formation	Argentina	Deseado Massif
Abanico Formation	Chile	Abanico Basin
Agua de la Piedra Formation	Argentina	Precordillera
Barzalosa Formation	Colombia	Upper Magdalena Valley
Castillo Formation	Venezuela	Falcón Basin
Chambira Formation	Peru	Ucayali Basin
Chilcatay Formation	Peru	Pisco Basin
Dos Bocas Formation	Ecuador	Progreso Basin
Fray Bentos Formation	Argentina Uruguay	Paraná Basin
Monte León Formation	Argentina	Austral Basin
Moquegua Formation	Peru	Moquegua Basin
Mugrosa Formation	Colombia	Middle Magdalena Valley
Petaca Formation	Bolivia	Subandean Belt
"Puca Group" - Lacayani fauna	Bolivia	Subandean Belt
Río Baguales Formation	Chile	Aysén Basin
Salla Formation	Bolivia	Salla-Luribay Basin
Sarmiento Formation	Argentina	Golfo San Jorge Basin
Tremembé Formation/São Paulo Formation	Brazil	Taubaté Basin

Fossils

Group	Fossils	Formation	Notes
Mammals	Ancylocoelus frequens, A. minor, Archaeohyrax patagonicus, Archaeutatus malaspinensis, Argyrohyrax proavus, Asmodeus osborni, Australohyaena antiqua, Cephalomys arcidens, C. plexus, Chubutomys simpsoni, Deseadomys arambourgi, Liarthrus copei, Litodontomys chubutensis, Meteutatus aff. lagenaformis, Parastrapotherium holmbergi, Progaleopithecus tournoueri, Protosteiromys asmeodeophilus, P. medianus, Prozaedyus aff. impressus, cf. Sallacyon hoffstetteri, Scotamys antiquus, Stenotatus aff. ornatus, Trachytherus spegazzinianus, Astrapotherium sp., Coresodon sp., Deuterotherium sp., Hegetotherium sp., Homalodontotherium sp., Leontinia sp., Nesodon sp., Octodontotherium sp., Trimerostephanos sp., Palaeothentidae indet.	Deseado
	Archaeohyrax suniensis, Argyrohyrax proavus, Asmodeus petrasnerus, Fieratherium sorex, Gualta cuyana, Hegetotheriopsis sulcatus, Mendozahippus fierensis, Meteutatus aff. lagenaformis, Proborhyaena gigantea, Prohegetotherium malalhuense, P. schiaffinoi, P. cf. sculptum, Propachyrucos cf. smithwoodwardi, ?Prozaedyus aff. impressus, Pyrotherium romeroi, Stenotatus aff. ornatus, Trachytherus cf. spegazzinianus, cf. Archaeotypotherium sp., Pharsophorus sp., Proadinotherium sp., Progaleopithecus sp., cf. Prosotherium sp., Acaremyidae indet., Glyptodontidae indet., Interatheriidae indet., Litopterna indet., ?Megalonychidae indet., Notohippidae indet., Toxodontidae indet.	Agua de la Piedra
	Lophiodolodus chaparralensis, Xenastrapotherium chaparralensis, Proadinotherium sp., Protheosodon sp., Astrapotheriidae indet.	Barzalosa
	Canaanimico amazonensis, Chambiramys shipiborum, C. sylvaticus, Deseadomys cf. arambourgi, Loretomys minutus, Palaeosteiromys amazonensis, Plesiosteiromys newelli, Maquiamys praecursor, Ucayalimys crassidens, Abderites sp., aff. Eosallamys sp., cf. Neoglyptatelus sp., Adelphomyinae indet., Anthropoidea indet., Astrapotheriidae indet., ?Caenolestidae indet., Caviomorpha indet., ?Chinchilloidea indet., Emballonuridae indet., ?Erethizontoidea indet., ?Herpetotheriidae indet., Interatheriinae indet, Litopterna indet., Marsupialia indet., Microbiotheria indet., Mylodontidae indet., Mylodontoidea indet., Notoungulata indet., Octodontoidea indet., ?Palaeothentidae indet., Pampatheriidae indet., ?Rhinolophoidea indet., Tolypeutinae indet., Toxodontidae indet., Typotheria indet., Vespertilionoidea indet.	Chambira
	Chilcacetus cavirhinus, Huaridelphis raimondii, Incacetus broggii, Macrosqualodelphis ukupachai, Notocetus vanbenedeni, cf. Kentriodon sp., Cetotheriidae indet., Eurhinodelphinidae indet., Mysticeti indet., Odontoceti indet., Pinnipedia indet., Physeteroidea indet., Squalodelphinidae indet.	Chilcatay
	Urkudelphis chawpipacha	Dos Bocas
	Dasypodon atavus, Eopachyrucos ranchoverdensis, Fiandraia romeii, Palaeopeltis inornatus, Palmiramys waltheri, Proborhyaena cf. gigantea, Prohegetotherium schiaffinoi, Proterotherium berroi, Pseudohegetotherium palmirense, Scarrittia robusta, Trachytherus spegazzinianus, Urotherium interundatum, Uruguaytherium beaulieui, Argyrohyrax sp., Berthawyleria sp., Cephalomyopsis sp., Doellotatus sp., Eoviscaccia sp., Hegetotherium sp., Protypotherium sp., ?Scelidodon sp., ?Vassallia sp., Archaeohyracidae indet., Dasypodidae indet., Euphractini indet., Glyptatelinae indet., Interatheriinae indet., Leontiniidae indet., Machlydotherium indet., Trachytheriinae indet.	Fray Bentos
	Odontoceti indet., Phocidae indet.	Gaiman
	Diaphorocetus poucheti, Notocetus vanbenedeni, Prosqualodon australis, cf. Kentriodon sp.	Monte León
	?Rhynchippus sp.	Petaca
	Eoviscaccia boliviana, Trachytherus spegazzinianus, Cephalomyopsis sp., Glyptatelus sp., Prohegetotherium sp., Borhyaenidae indet., Dasypodidae indet., Macraucheniidae indet., Orophodontidae indet.	Lacayani fauna
	Anayatherium ekacoa, A. fortis, Andinogale sallensis, Antawallathentes illimani, A. quimsacruza, Archaeohyrax suniensis, Branisamys luribayensis, Branisella boliviana, Brucemacfaddenia boliviensis, Cephalomys bolivianus, ?Coniopternium primitivum, Eurygenium pacegnum, Evolestes hadrommatos, Federicoanaya sallaensis, Incamys bolivianus, Kuntinaru boliviensis, Luribayomys masticator, Migraveramus beatus, Notogale mitis, Palaeothentes boliviensis, Paraborhyaena boliviana, Paroctodontotherium calleorum, Pascualihippus boliviensis, Pharsophorus lacerans, Proargyrolagus bolivianus, Prohegetotherium schiaffinoi, Pseudoglyptodon sallaensis, Pyrotherium macfaddeni, cf. Pyrotherium romeroi, Ronwolffia pacifica, Rhynchippus cf. brasiliensis, Sallacyon hoffstetteri, Salladolodus deuterotherioides, Sallamys pascuali, Sallatherium altiplanense, Thadanius hoffstetteri, Trachytherus alloxus, Tricoelodus boliviensis, cf. Acamana sp., cf. Eocoleophorus sp., Glyptatelus sp., aff. Neoreomys sp., Peltephilus sp., Astrapotheriidae indet., Borhyaeninae indet., Dasypodidae indet., Dasyproctidae indet., Megalonychidae indet., Nesodontinae indet., Polydolopidae indet.	Salla
	Acarechimys leucotheae, Ancylocoelus frequens, Archaeohyrax patagonicus, Archaeophylus patrius, Archaeotypotherium propheticus, Archaeutatus malaspinensis, Argyrohyrax proavus, Asmodeus osborni, Asteromys punctus, Australohyaena antiqua, Bryanpattersonia sulcidens, Cephalomys arcidens, C. ceciae, C. plexus, Changquin woodi, Chubutomys navaensis, C. simpsoni, Clypeotherium magnum, Coresodon scalpridens, Cramauchenia normalis, Deseadomys arambourgi, Deuterotherium distichum, Draconomys verai, Elmerriggsia fieldia, Eomicrobiotherium matutinum, Eopachyrucos pliciformis, ?Eoviscaccia australis, Epiklohnia verticalis, Ethelomys loomisi, Eurygenium latirostris, Eutrachytherus grandis, Galileomys baios, Hegetotheriopsis sulcatus, Henricofilholia lustrata, H. vucetichia, Hondonadia parca, H. cf. pumila, Incamys bolivianus, I. menniorum, Kramadolops maximus, Leontinia gaudryi, Leucokephalos zeffiae, Liarthrus copei, Litodontomys chubutensis, Llitun notuca, Loncolicu tretos, Maddenia lapidaria, Medistylus dorsatus, Meteutatus lagenaformis, Notodiaphorus crassus, Notogale mitis, Palaeopeltis inornatus, Palaeothentes chubutensis, P. lucina, P. praecursor, Parabderites minusculus, Parastrapotherium holmbergi, Parutaetus ?chilensis, Patagonhippus canterensis, P. dukei, Pharsophorus lacerans, P. tenuis, Pilchenia intermedia, Platypittamys brachyodon, Proadiantus excavatus, P. gibbus, Proadinotherium leptognathus, Proborhyaena gigantea, Proeuphractus setiger, Progaleopithecus fissurellatus, P. tournoueri, Prohegetotherium sculptum, Propachyrucos ameghinorum, Prosotherium garzoni, P. triangulidens, ?Protacaremys adilos, Protarchaeohyrax minor, Protheosodon coniferus, Protosteiromys asmodeophilus, P. medianus, Prozaedius impressus, P. planus, Pseudhalmarhiphus guaraniticus, Pyrotherium sorondoi, Rhynchippus equinus, R. pumilus, Rosendolops cf. ebaios, Sadypus tortuosus, Santiagorothia chiliensis, Scarrittia barranquensis, S. canquelensis, Scotamys antiquus, ?Stenotatus aff. ornatus, Trachytherus spegazzinianus, Tricoelodus bicuspidatus, Vallehermosomys mazzonii, ?V. merlinae, Barrancatatus sp., Cephalomyopsis sp., Cochilius sp., Evolestes sp., Octodontotherium sp., Peltephilus sp., Prohegetotherium sp., Trachytherus sp., Caviomorpha indet., ?Chinchilloidea indet., ?Dasyproctidae indet., cf. Eobranisamys indet., Eocardiidae indet., Henricosborniidae indet., Interatheriinae indet., Isotemnidae indet., Macraucheniidae indet., Palaeothentidae indet., Pichipilidae indet., Propalaehoplophorinae indet., Rodentia indet., Stegotheriini indet., Toxodontia indet.	Sarmiento
	cf. Leontinia gaudryi	São Paulo
	Moqueguahippus glycisma, Rhynchippus cf. pumilus, Sallamys quispea, Trachytherus ramirezi, T. cf. spegazzinianus, cf. Coniopternium sp., cf. Dasypodinae indet., Hegetotheriidae indet., Macraucheniidae indet., Proterotheriidae indet.	Moquegua
Birds	Cladornis pachypus, Cruschedula revola	Deseado
	cf. Andrewsornis sp.	Agua de la Piedra
	Palaeospheniscus sp.	Chilcatay
	Devincenzia gallinali	Fray Bentos
	Eretiscus tonnii, Palaeospheniscus patagonicus, Palaeospheniscus wimani, Paraptenodytes antarcticus	Gaiman
	Arthrodytes andrewsi, Paraptenodytes brodkorbi, P. robustus, Pseudospheniscus concavus	Monte León
	Spheniscidae indet.	Río Baguales
	Phorusrhacidae indet.	Salla
	Physornis sp.	Sarmiento
	Paraphysornis brasiliensis, Taubatornis campbelli	Tremembé
	Phorusrhacidae indet.	Moquegua
Reptiles & amphibians	Balanerodus logimus, Gavialidae indet.	Barzalosa
	?Balanerodus sp., Podocnemis sp., cf. Purussaurus sp., cf. Sebecus sp., Anura indet., Booidea indet., Caimaninae indet., Colubroidea indet., Gavialoidea indet., Serpentes indet.	Chambira
	Testudines indet.	Chilcatay
	Testudinidae indet.	Fray Bentos
	Eusuchia indet., Sebecosuchia indet.	Mugrosa
	?Chelonoidis sp.	Petaca
	cf. Calyptocephalella sp., Testudines indet.	Salla
	Calyptocephalella canqueli, Neoprocoela edentatus, Eupsophus sp., ?Madtsoia sp., Iguaninae indet., Sebecidae indet.	Sarmiento
	Caiman tremembensis	Tremembé
Fishes	cf. Hydrolycus sp., Leporinus sp., cf. Phractocephalus sp., Potamotrygon sp., Actinopterygii indet., Cichlidae indet., ?Erythrinidae indet., Loricariidae indet., Pimelodidae indet.	Chambira
	Carcharhinus cf. brachurus, Carcharodon hastalis, Carcharodon subauriculatus, Hemipristis cf. serra, Isurus desori	Chilcatay
	Carcharodon hastalis, Carcharoides catticus, Echinorhinus pozzii, Galeocerdo aduncus, Lamna totuserrata, Megascyliorhinus trelewensis, Heterodontus sp., Labrodon sp., Sphyrna sp., Squatina sp., Molidae indet., Myliobatoidea indet.	Gaiman
	Isurus dolloi, Striatolamia macrota, Abdounia sp.	Río Baguales
	Taubateia paraiba	Tremembé
Insects	?Archaeodrapetiops elongata, A. mezzalirai, A. nefera, A. transversa, Archaeolycorea ferreirai, Eternia papaveroi, Indusia suguioi, Kleopathra nemogypsia, K. noctodiva, Microbasis longinota, Neorinella garciae, Parafitopteryx duarteae, Paratilgidopsis praecursora, Petisca dryellina, Philodarchia cigana, Psephenella ferreirai, ?Tabanus tremembeensis, Taubocicadellina breviptera, Taubarixa macrocelata, Tauborixella santosae, Tauborixiellopsis breviclavata, Taubatehymen minuta, Tremembaetalion minutum, Tremembecarabus rotundus, Tremembella gracilis, Tremembellina microcelata, Trulaxia primula, Curculioninae indet., Hydrophilidae indet.	Tremembé

Correlations

The Deseadan South American land mammal age (SALMA) is equivalent to the Arikareean in the North American land mammal age (NALMA) and the Harrisonian in the 2000 version of the classification. It overlaps with the Hsandagolian of Asia and the MP 25 zone of Europe, the Waitakian and the Landon epoch of New Zealand.

Deseadan correlations in South America

Formation	Agua	Rancahué	Guillermo	Deseado	Sarmiento	Salla	Lacayani	Fray Bentos	Moquegua	Chambira	Barzalosa	Tremembé	Cascadas	Map
Basin	Neuquén		Austral	Deseado	San Jorge	Salla	Subandean	Norte	Moquegua	Ucayali	VSM	Taubaté	Panama	Deseadan (South America)
Country	Argentina					Bolivia		Uruguay	Peru		Colombia	Brazil	Panama
Archaeohyrax
Prohegetotherium
Pyrotherium
Pharsophorus
Trachytherus
Proadinotherium
Proborhyaena
Meteutatus
Andrewsornis
Terror birds
Rodents
Reptiles
Primates
Flora
Insects
Environments	Alluvial		Fluvial	Fluvial		Alluvial	Fluvial-alluvial	Fluvial	Fluvio-lacustrine		Alluvial-fluvial	Lacustrine	Fluvial	Deseadan volcanoclastics Deseadan fauna Deseadan flora
Volcanic	Yes				Yes	Yes	Yes		Yes				Yes

References

Bibliography

General

• Woodburne, M.O.; Goin, F.J.; Bond, M.; Carlini, A.A.; Gelfo, J.N.; López, G.M.; Iglesias, A.; Zimicz, A.N. (2013), "Paleogene Land Mammal Faunas of South America; a Response to Global Climatic Changes and Indigenous Floral Diversity", Journal of Mammalian Evolution, 21: 1–73, doi:10.1007/s10914-012-9222-1, hdl:11336/31253, retrieved 2019-02-15

Deseado Formation

• Agnolin, Federico (2006), "Posición sistemática de algunas aves fororracoideas (Ralliformes: Cariamae) Argentinas", Revista del Museo de Ciencias Naturales, 8: 27–33, doi:10.22179/REVMACN.8.348, retrieved 2018-09-10
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Abanico Formation

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Agua de la Piedra Formation

• Cerdeño, Esperanza; Reguero, Marcelo (2015), "The Hegetotheriidae (Mammalia, Notoungulata) assemblage from the late Oligocene of Mendoza, central-western Argentina", Journal of Vertebrate Paleontology, 35 (2) e907173, Bibcode:2015JVPal..35E7173C, doi:10.1080/02724634.2014.907173, hdl:11336/48566, retrieved 2015-02-13
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Barzalosa Formation

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Castillo Formation

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Chambira Formation

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Chilcatay Formation

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Dos Bocas Formation

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Fray Bentos Formation

• Bond, Mariano; López, Guillermo; Reguero, Marcelo A.; Scillato Yané, Gustavo J.; Vucetich, María G. (1998), "Los mamíferos de la Formación Fray Bentos (Edad mamífero Deseadense, Oligoceno Superior) de las provincias de Corrientes y Entre Ríos, Argentina", Asociación Paleontológica Argentina. Publicación Especial, 5: 41–50, retrieved 2018-09-12
• Tófalo, Ofelia Rita; Morrás, Héctor J.M. (2009), "Evidencias paleoclimáticas en duricostras, paleosuelos y sedimentitas silicoclásticas, del Cenozoico de Uruguay" (PDF), Revista de la Asociación Geológica Argentina: 674–686, retrieved 2018-09-12

Gaiman Formation

• Acosta Hospitaleche, Carolina (2007), "Revisión sistemática de Palaeospheniscus biloculata (Simpson) nov. comb. (Aves, Spheniscidae) de la Formación Gaiman (Mioceno Temprano), Chubut, Argentina", Ameghiniana, 44: 417–426, retrieved 2018-09-11
• Viglino, Mariana; Buono, Mónica R.; Gutstein, Carolina S.; Cozzuol, Mario A.; Cuitiño, José I. (2018), "A new dolphin from the early Miocene of Patagonia, Argentina: Insights into the evolution of Platanistoidea in the Southern Hemisphere" (PDF), Acta Palaeontologica Polonica, 63 (2): 261–277, doi:10.4202/app.00441.2017, retrieved 2019-02-11

Lacayani fauna

• Billet, Guillaume; De Muizon, Christian; Mamani Quispe, Bernardino (2008), "Late Oligocene mesotheriids (Mammalia, Notoungulata) from Salla and Lacayani (Bolivia): implications for basal mesotheriid phylogeny and distribution", Zoological Journal of the Linnean Society, 152: 153–200, doi:10.1111/j.1096-3642.2007.00388.x, retrieved 2018-09-11

Monte León Formation

• Náñez, Carolina; Quattrocchio, Mirta E.; Ruiz, Liliana (2009), "Palinología y micropaleontología de las Formaciones San Julián y Monte León (Oligoceno - Mioceno temprano) en el subsuelo de cabo Curioso, provincia de Santa Cruz, Argentina", Ameghiniana, 46: 669–693, retrieved 2017-08-15

Moquegua Formation

• Alván, Aldo; Ramírez, Katerin; von Eynatten, Hilmar; Dunkl, István; Jacay, Javier; Bertone, Gabriela (2017), "Evolución Geológica de las Cuencas de Antearco del Sur de Perú (Moquegua y Camaná-Mollendo): Proveniencia Sedimentaria y Análisis de Facies en Rocas Cenozoicas", Boletín de la Sociedad Geológica del Perú, 112: 53–77, retrieved 2018-09-10
• Shockey, Bruce J.; Billet, Guillaume; Salas Gismondi, Rodolfo (2016), "A new species of Trachytherus (Notoungulata: Mesotheriidae) from the late Oligocene (Deseadan) of Southern Peru and the middle latitude diversification of early diverging mesotheriids", Zootaxa, 4111 (5): 565–583, doi:10.11646/zootaxa.4111.5.3, PMID 27395103

Mugrosa Formation

• Gómez, Luis Alfonso; Patiño, Alejandro; Renzoni, Giancarlo; Beltrán, Alejandro; Quintero, Claudia; Manrique, Martín (2008), Geología de la Plancha 119 Barrancabermeja - 1:100,000, INGEOMINAS, pp. 1–105

Petaca Formation

• Hernández, R.; Jordan, T.; Dalenz Farjat, A.; Echavarría, L.; Idelman, B.; Reynolds, J. (2002), Edades, distribución, controles tectónicos y eustáticos de las ingresiones marinas paranense y caribeana en el sur de Bolivia y Argentina (PDF), V Congreso Latinoamericano de Paleontología, pp. 1–27, retrieved 2018-09-10

Río Baguales Formation

• Le Roux, Jacobus P.; Puratich, Jacqueline; Mourgues, F. Amaro; Oyarzún, José Luis; Otero, Rodrigo A.; Torres, Teresa; Hervé, Francisco (2010), "Estuary deposits in the Río Baguales Formation (Chattian-Aquitanean), Magallanes Province, Chile" (PDF), Andean Geology, 37: 329–344, retrieved 2017-10-25

Salla Formation

• Reguero, Marcelo A.; Cerdeño, Esperanza (2005), "New late Oligocene Hegetotheriidae (Mammalia, Notoungulata) from Salla, Bolivia", Journal of Vertebrate Paleontology, 25 (3): 674–684, doi:10.1671/0272-4634(2005)025[0674:NLOHMN]2.0.CO;2, retrieved 2018-09-11
• Rincón, Ascanio D.; Shockey, Bruce J.; Anaya, Federico; Solórzano, Andrés (2015), "Palaeothentid Marsupials of the Salla Beds of Bolivia (Late Oligocene): Two New Species and Insights into the Post-Eocene Radiation of Palaeothentoids", Journal of Mammalian Evolution, 22 (4): 455–471, doi:10.1007/s10914-015-9295-8, retrieved 2019-02-13
• Shockey, Bruce J (2017), "New early diverging cingulate (Xenarthra: Peltephilidae) from the Late Oligocene of Bolivia and considerations regarding the origin of crown Xenarthra", Bulletin of the Peabody Museum of Natural History, 58 (2): 371–396, retrieved 2019-02-12

Sarmiento Formation

• Acosta Hospitaleche, Carolina; Tambussi, Claudia (2005), "Phorusrhacidae Psilopterinae (Aves) en la Formación Sarmiento de la localidad de Gran Hondonada (Eoceno Superior), Patagonia, Argentina [Phorusrhacidae Psilopterinae (Birds) in the Sarmiento Formation from the Gran Hondonada locality (Upper Eocene), Patagonia, Argentina]" (PDF), Revista Española de Paleontología, 20: 127–132, retrieved 2018-09-10
• Arnal, Michelle; Kramarz, Alejandro G.; Vucetich, M. Guiomar; Vieytes, E. Carolina (2014), "A new early Miocene octodontoid rodent (Hystricognathi, Caviomorpha) from Patagonia (Argentina) and a reassessment of the early evolution of Octodontoidea", Journal of Vertebrate Paleontology, 34 (2): 397–406, Bibcode:2014JVPal..34..397A, doi:10.1080/02724634.2013.808203, hdl:11336/29748, retrieved 2019-02-12
• Cheme Arriaga, Lucas; Dozo, María Teresa; Gelfo, Javier N. (2016), "A new Cramaucheniinae (Litopterna, Macraucheniidae) from the early Miocene of Patagonia, Argentina", Journal of Vertebrate Paleontology, 36 (6) e1229672, Bibcode:2016JVPal..36E9672C, doi:10.1080/02724634.2017.1229672, hdl:11336/30386, retrieved 2019-02-12
• Dozo, María Teresa; Ciancio, Martín; Bouza, Pablo; Martínez, Gastón (2014), "Nueva asociación de mamíferos del Paleógeno en el este de la Patagonia (provincia de Chubut, Argentina): implicancias biocronológicas y paleobiogeográficas" (PDF), Andean Geology, 41: 224–247, Bibcode:2014AndGe..41a...9D, doi:10.5027/andgeoV41n1-a09, retrieved 2017-08-15
• Pérez, María Encarnación; Krause, Marcelo; Vucetich, María Guiomar (2012), "A new species of Chubutomys (Rodentia, Hystricognathi) from the late Oligocene of Patagonia and its implications on the early evolutionary history of Cavioidea sensu stricto", Geobios, 45 (6): 573–580, Bibcode:2012Geobi..45..573P, doi:10.1016/j.geobios.2012.06.001
• Shockey, Bruce J.; Flynn, John J.; Croft, Darin A.; Gans, Phillip; Wyss, André R. (2012), "New leontiniid Notoungulata (Mammalia) from Chile and Argentina: comparative anatomy, character analysis, and phylogenetic hypotheses", American Museum Novitates (3737): 1–64, doi:10.1206/3737.2, retrieved 2019-02-15
• Sterli, Juliana; De la Fuente, Marcelo S.; Krause, J. Marcelo (2015), "A new turtle from the Palaeogene of Patagonia (Argentina) sheds new light on the diversity and evolution of the bizarre clade of horned turtles (Meiolaniidae, Testudinata)", Zoological Journal of the Linnean Society, 174 (3): 519–548, doi:10.1111/zoj.12252, hdl:11336/41594, retrieved 2019-02-13
• Vucetich, M.G.; Dozo, M.T.; Arnal, M.; Pérez, M.E. (2015), "New rodents (Mammalia) from the late Oligocene of Cabeza Blanca (Chubut) and the first rodent radiation in Patagonia", Historical Biology: An International Journal of Paleobiology, 27 (2): 236–257, Bibcode:2015HBio...27..236V, doi:10.1080/08912963.2014.883506, hdl:11336/17958, retrieved 2019-02-13
• Vucetich, María G.; Pérez, María E.; Ciancio, Martín R.; Carlini, Alfredo A.; Madden, Richard H.; Kohn, Matthew J. (2014), "A new acaremyid rodent (Caviomorpha, Octodontoidea) from Scarritt Pocket, Deseadan (late Oligocene) of Patagonia (Argentina)", Journal of Vertebrate Paleontology, 34 (3): 689–698, Bibcode:2014JVPal..34..689V, doi:10.1080/02724634.2013.804414, hdl:11336/17973, retrieved 2019-02-12
• Wyss, André R.; Flynn, John J.; Croft, Darin A. (2018), "New Paleogene notohippids and leontiniids (Toxodontia; Notoungulata; Mammalia) from the Early Oligocene Tinguiririca Fauna of the Andean Main Range, central Chile", Journal of South American Earth Sciences, 3903: 1–42, hdl:2246/6901, retrieved 2019-02-11

Tremembé Formation

• Do Couto Ribeiro, Graziella (2010), Avaliação morfológica, taxonômica e cronológica dos mamíferos fósseis da Formação Tremembé (Bacia de Taubaté), Estado de São Paulo, Brasil, Universidade de São Paulo, pp. 1–112
• Malabarba, Maria Claudia; Lundberg, John G. (2007), "A fossil loricariid catfish (Siluriformes: Loricarioidea) from the Taubaté Basin, eastern Brazil", Neotropical Ichthyology, 5 (3): 263–270, doi:10.1590/S1679-62252007000300005, retrieved 2018-10-08
• Olson, Storrs L.; Alvarenga, Herculano M.F. (2002), "A new genus of small teratorn from the Middle Tertiary of the Taubate Basin, Brazil (Aves: Teratornithidae)" (PDF), Proceedings of the Biological Society of Washington, 115: 701–705, retrieved 2017-09-04